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A demesne is a sub-national division of Natopia. The term "demesne" has been in near-continuous use to describe Natopia's subdivisions since 2005. Although originally intended to imply direct feudal subservience to the central government, the autonomy granted to demesnes of Natopia have fluctuated over the years. Customs, traditions, symbols, cities, and general cultural development have been the primary domain of the demesnes. The imperial government, except in extreme circumstances, allows the demesnes to govern themselves within the limits of the Caprine Code.

The current arrangement of demesnes includes a relatively new category: the demesne union. Demesne Unions generally consist of political, dynastic, geographic, or religious groupings of demesnes. Some demesne unions effectively function as a single demesne, albeit divided into smaller demesnes for historic, demographic, or cultural purposes, (e.g. The Most Serene Union, the Elian Union). Other demesnes are more feudal in nature, with one dominant demesne holding control over 'junior' demesnes (e.g. Flaventia). Other demesne unions merely act as the legal mechanism to unite demesnes under a single government that exists above the governments of the demesnes (e.g. Uppland and Arboria). All demesnes, regardless of its status within a demesne union, send representatives to the Frenzy.

Demesnes of Natopia